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Originally Posted by Iuris
Essentially, it would appear that inter-species tension existed for a long time.
However, the Titans eventually subverted the Charr into their servants, supplied them with destructive magics that brought down the wall, and conquered Tyria. The Charr went for the rest of humanity too, but were stopped. In the case of Kryta, by the White mantle with their Mursaat lords and in the case of Orr, by a suicidal spell that took out both the Charr and Orr.
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Yep, do the quest (or read the quest dialog from Guildwiki) "A Flickering Flame" and and the unknown reasons other posters talked about become a little less unknown. As I recall there was some other dialog in pop-up speech boxes in the quest "Dismember the Titans" that expand on it some more, but the wiki doesn't list it - it may be another quest.
It had to do with the destruction of Arah, Terick used the Charr invasion to push the Vizier into incanting the magic that destroyed Orr (and the City of the Gods, Arah). See the quests "Escape from the Torment" and "Blueprint of the Fall". Effectively Abbadon was influencing both races (Charr and Humans) into a situation where Arah was destroyed.
Of course, there is also those pesky Flameseeker prophecies in there - I rather suspect that Arah was just just a nice side prize. But that isn't exactly what was being asked - the Charr were manipulated by the titans who were servants of Abbadon into invading the human world. If the main goal was the destruction of Arah or the Flameseeker prophecies is unknown. As far as I know there is in game dialog that suggests both.
Of course, this is also why so many say GW story sucks - they either do not do the quests, do not read them, or simply do not remember. Unlike many other traditional RPG's, GW doesn't force you to read the story.