Two people in particular sparked the thought that brings this theory about, one that the Mursaat were the "architects" of the world, the other that the eye itself is actually a representation of the Five Gods. I think that, in fact, the eye is actually a representation of the Five Gods, that it is a symbol of them watching over Tyria. However, why place such symbols of importance in these strange isolated locations if you want your worshipers to know of it? This is where the Mursaat come in, I think that they worked in the background, building structures and cities before any human. Worshiping the Five Gods at one time themselves, and followed in their every step.
When the Exodus occurred, they felt betrayed, and abandoned their Temples, but did not deny their existence themselves. How could they? They sat on proof in the Ring of Fire Islands Chain, and in Bloodstone Fen. However, they needed to keep the Five Gods' remnants contained, their mess, so to speak. The Bloodstones were mainly guarded by them and they chose to use the ones that were close enough to be of use. It would seem likely that they had not been sacrificing the Chosen prior to their meeting of Saul, and it was by the Eye of Janthir that they discovered this property of the humans. That there was something significant about them, they must have discovered eventually that their blood could activate the Bloodstones.
I suspect that the runes and glyphs we see on Bloodstone Fen and the Bloodstone in Hell's Precipice were not actually there originally, rather they were put there by the Mursaat. Take note that the Bloodstone in the Bloodstone Caves shows no such markings and it looks very much like what may be a Bloodstone in its primitive state.
However, it also seems that the Mursaat may have known of its presence, and learned of another use for the runes that they had used previously. As seen here:
However, I have overlooked something, return to the Temples for a moment and then remember the Temple of Grenth in the Underworld, it seems possible that the Five Gods did not notice their devotion. As such they most likely contracted them to build Temples of grand magnificence in places chosen by the Gods themselves. Grenth perhaps even allowed their presence in the Underworld to construct his very own palace, his throne, of the Underworld.
What brings me to this conclusion is something that has caused endless debate amongst those researching here.
This:
And this:
I think that I, with ideas from one who suggested they were architects, have found the link here. However, you may ask, what does this have to do with Abaddon's taste for eyes? The eyes are a symbol of the Five Gods, as I stated earlier, the Gods erased nearly every trace of Abaddon. His revenge is taken in a similar way, in feasting on the eyes of those who worship the Five Gods. Their symbol becomes torment for their devout worshipers.
Another element, to add to the Mursaat being architects, is found in Abaddon's domain, actually. Something I had noticed quite often as I explored the Atrocity Library is the bridge at the entrance to the Domain of Secrets. Its architecture resembles the Temples to Grenth, and Balthazar, and the very architecture of Grenth's own palace. What secrets are there to be found in that? The Mursaat while devout worshipers, did not want to be known by the other races, indeed it was only by chance Saul found them. So in a way it represents the secrets Abaddon held about the origins of this strange, yet magnificent architecture.
